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Corresponding author is indicated by asterisk (*)


  • Chao A.*, Kubota Y., Zelený D., Chiu C.-H., Li C.-F., Kusumoto B., Yasuhara M., Thorn S., Wei C.-L., Costello M.J. & Colwell R.K. (2020): Quantifying sample completeness and comparing diversities among assemblages. Ecological Research, 35, 292–314 (www & pdf)
  • Chudomelová M.* & Zelený D. (2020): Tracing the signs of local dispersal in the temperate forest understorey using spatially structured vegetation data. Journal of Vegetation Science, 31, 84–94 (www, read-only pdf, Behind the paper post on


  • Kattge J., Bönisch G., Díaz S., Lavorel S., Prentice I. C., Leadley P., Tautenhahn S., Gijsbert D., … Zelený D., … Wirth C. (2019): TRY plant trait database – enhanced coverage and open access. Global Change Biology, 26, 119–188 (www & pdf)
  • Zelený D. & Chytrý M. (2019): Ecological Specialization Indices for species of the Czech flora. Preslia 91: 93–-116 (pdf, Appendix 1, Supplementary online information on GitHub)
  • Zelený D. & Fidelis, A. (2019): About travel awards to annual symposium by IAVS Global Sponsorship Committee. IAVS Bulletin, 4, 14–16 (pdf)
  • Wiser, S., Zelený, D., Diekmann, M., Fidelis, A., Janišová, M., Loidi, J. & Minchin, P. (2019): Meet the new IAVS Governing Board. IAVS Bulletin, 4, 2–13 (pdf)


  • Zelený D. (2018): Which results of the standard test for community weighted mean approach are too optimistic? Journal of Vegetation Science 29: 953-966 (www, read-only pdf, preprint on BioRxiv, plain language summary on, blog post on
  • Chytrý M., Tichý L., Dřevojan P., Sádlo J. & Zelený D. (2018): Ellenberg-type indicator values for the Czech flora. Preslia 90: 83–103 (www, pdf)


  • Zelený D. (2017): Bias in community-weighted mean analysis of plant functional traits and species indicator values. bioRxiv. DOI: 10.1101/046946 (see also my blog post)
  • Bonari, G., Fajmon, K., Malenovský, I., Zelený, D., Holuša, J., Jongepierová, I., Kočárek, P., Konvička, O., Uřičář, J. & Chytrý, M. (2017): Management of semi-natural grasslands benefiting both plant and insect diversity: The importance of heterogeneity and tradition. Agriculture, Ecosystems & Environment 246: 243-252 (www)
  • Wagner V., Chytrý M., Zelený D., von Wehrden H., Brinkert A., Danihelka J., Jansen F., Hölzel N., Kamp J., Lustyk P., Merunková K., Palpurina S., Preislerová Z. & Wesche K. (2017): Regional differences in soil pH niche among dry grassland plants in Eurasia. Oikos, 126: 660–670 (www)
  • Hawkins B.A., Leroy B., Rodríguez M.Á., Singer A., Vilela B., Villalobos F., Wang X. & Zelený D. (2017): Structural bias in aggregated trait variables driven by species co-occurrences: a pervasive problem in community and assemblage data. Journal of Biogeography, 44: 1199–1211 (www)
  • Chudomelová M., Zelený D. & Li C.-F. (2017): Contrasting patterns of fine-scale herb layer species composition in temperate forests. Acta Oecologica, 80: 24-31 (www)


  • Divíšek J., Storch D., Zelený D. & Culek M. (2016): Towards the spatial coherence of biogeographical regionalizations at sub-continental and landscape scales. Journal of Biogeography, 43: 2489–2501 (www)
  • de Bello F., Fibich P., Zelený D., Kopecký M., Mudrák O., Chytrý M., Pyšek P., Wild J., Michalcová D., Sádlo J., Šmilauer P., Lepš J. & Pärtel M. (2016): Measuring size and composition of species pools: a comparison of dark diversity estimates. Ecology and Evolution, 6: 4088–4101 (www)
  • Carboni M., Zelený D. & Acosta A. (2016): Measuring ecological specialization along a natural stress gradient using a set of complementary niche breadth indices. Journal of Vegetation Science, 27: 892-903 (www)


  • Chytrý M., Zelený D. & Hettenbergerová E. (eds.) (2015): 58th Annual Symposium of the International Association for Vegetation Science: Understanding broad-scale vegetation patterns. 19–24 July 2015, Brno, Czech Republic. Abstracts. Masaryk University, Brno. (pdf)
  • Li C.-F., Zelený D., Chytrý M., Chen M.-Y., Chen T.-Y., Chiou C.-R., Hsia Y.-J., Liu H.-Y., Yang S.-Z., Yeh C.-L., Wang J.-C., Yu C.-F., Lai Y.-J., Guo K. & Hsieh C.-F. (2015): Chamaecyparis mountain cloud forest in Taiwan: ecology and vegetation classification. Ecological Research, 30: 771-791 (www, pdf)
  • Lososová Z., de Bello F., Chytrý M., Kühn I., Pyšek P., Sádlo J., Winter M. & Zelený D. (2015): Alien plants invade more phylogenetically clustered community types and cause even stronger clustering. Global Ecology and Biogeography, 24: 786–794 (www)
  • Čeplová N., Lososová Z., Zelený D., Chytrý M., Danihelka J., Fajmon K., Láníková D., Preislerová Z., Řehořek V. & Tichý L. (2015): Phylogenetic diversity of central European urban plant communities: effects of alien species and habitat types. Preslia 87: 1–16 (pdf)


  • Horsák M., Zelený D. & Hájek M. (2014): Land snail richness and abundance along a sharp ecological gradient at two sampling scales: disentangling relationships. Journal of Molluscan Studies, 80: 256–264 (www)
  • Divíšek J., Zelený D., Culek M. & Šťastný K. (2014): Natural habitats matter: determinants of spatial pattern in animal assemblages of the Czech Republic. Acta Oecologica, 59: 7–17 (www)
  • Michalcová D., Chytrý M., Pechanec V., Hájek O., Jongepier J.W., Danihelka J., Grulich V., Šumberová K., Preislerová Z., Ghisla A., Bacaro B. & Zelený D. (2014): High plant diversity of grasslands in the landscape context: a comparison of contrasting regions in Central Europe. Folia Geobotanica, 49: 117–135 (www)


  • Chytrý M., Zelený D., Navrátilová J. & Sádlo J. (2013): Boreokontinentální jehličnaté lesy (Vaccinio-Picetea). Boreo-continental coniferous forests (Vaccinio-Picetea). In.: Chytrý M. (ed.), Vegetace České Republiky 4. Lesní a křovinná vegetace. Vegetation of the Czech Republic 4. Forest and Scrub Vegetation. Academia, Praha. pp. 380–432 (project website, Academia website)
  • Hettenbergerová E., Horsák M., Chandran R., Hájek M., Zelený D. & Dvořáková J. (2013): Patterns of land snail assemblages along a fine-scale moisture gradient. Malacologia 56: 31–42 (www)
  • Chang L.-W., Zelený D., Li C.-F., Chiu S.-T. & Hsieh C.-F. (2013): Better environmental data may reverse conclusions about niche- and dispersal-based processes in community assembly. Ecology 94: 2145–2151 (pdf)
  • Li C.-F., Chytrý M., Zelený D., Chen M.-Y., Chen T.-Y., Chiou C.-R., Hsia Y.-J., Liu H.-Y., Yang S.-Z., Yeh C.-L., Wang J.-C., Yu C.-F., Lai Y.-J., Chao W.-C. & Hsieh C.-F. (2013): Classification of Taiwan forest vegetation. Applied Vegetation Science, 16: 698–719 (www, related website with CoDeK application)
  • Hettenbergerová E., Hájek M., Zelený D., Jiroušková J. & Mikulášková E. (2013): Changes in species richness and species composition of vascular plants and bryophytes along a moisture gradient. Preslia, 85: 369–388 (pdf)
  • Fajmonová Z., Zelený D., Syrovátka V., Vončina G. & Hájek M. (2013): Distribution of habitat specialists in semi-natural grasslands. Journal of Vegetation Science, 24: 616–627 (www)


  • Lin C.-T., Li C.-F., Zelený D., Chytrý M., Nakamura Y., Chen M.-Y., Chen T.-Y., Hsia Y.-J., Hsieh C.-F., Liu H.-Y., Wang J.-C., Yang S.-Z., Yeh C.-L. & Chiou C.-R. (2012): Classification of the high-mountain coniferous forests in Taiwan. Folia Geobotanica, 47: 373–401 (www)
  • Axmanová I., Tichý L., Fajmonová Z., Hájková P., Hettenbergerová E., Li C.-F., Merunková K., Nejezchlebová M., Otýpková Z., Vymazalová M. & Zelený D. (2012): Estimation of herbaceous biomass from species composition and cover. Applied Vegetation Science, 15: 580–589 (www)
  • Zelený D. (2012): Poznámky k používání průměrných Ellenbergových indikačních hodnot při analýze vegetačních dat [Notes to the use of mean Ellenberg indicator values in vegetation analyses]. Zprávy České Botanické Společnosti, 47: 159–178 (pdf, related website)
  • Chytrý M., Ermakov N., Danihelka J., Hájek M., Hájková P., Horsák M., Kočí M., Kubešová S., Lustyk P., Otýpková Z., Pelánková B., Valachovič M. & Zelený D. (2012): High species richness in hemiboreal forests of the northern Russian Altai, southern Siberia. Journal of Vegetation Science, 23:605–616 (www)
  • Axmanová I., Chytrý M., Zelený D., Li C.-F., Vymazalová M., Danihelka J., Horsák M., Kočí M., Kubešová S., Lososová Z., Otýpková Z., Tichý L., Martynenko V.B., Baisheva E., Schuster B. & Diekmann M. (2012): The species richness–productivity relationship in the herb layer of European deciduous forests. Global Ecology and Biogeography, 21: 657–667 (www)
  • Zelený D. & Schaffers A.P. (2012): Too good to be true: pitfalls of using mean Ellenberg indicator values in vegetation analyses. Journal of Vegetation Science, 23: 419–431 (www, read-only pdf, software for modified permutation test, blog post)


  • Axmanová I., Zelený D., Li C.-F. & Chytrý M. (2011): Environmental factors influencing herb layer productivity in Central European oak forests: insights from soil and biomass analyses and a phytometer experiment. Plant and Soil, 342: 183–194 (www)


  • Chytrý M., Danihelka J., Axmanová I., Božková J., Hettenbergerová E., Li C.-F., Rozbrojová Z., Sekulová L., Tichý L., Vymazalová M. & Zelený D. (2010): Floristic diversity of an eastern Mediterranean dwarf shrubland: the importance of soil pH. Journal of Vegetation Science, 21: 1125–1137 (www)
  • Zelený D., Li C.-F. & Chytrý M. (2010): Pattern of local plant species richness along a gradient of landscape topographical heterogeneity: result of spatial mass effect or environmental shift? Ecography, 33:578–589 (pdf)
  • Tichý L., Hájek M. & Zelený D. (2010): Imputation of environmental variables for vegetation plots based on compositional similarity. Journal of Vegetation Science, 21: 88–95 (www)


  • Roleček J., Tichý L., Zelený D. & Chytrý M. (2009): Modified TWINSPAN classification in which the hierarchy respects cluster heterogeneity. Journal of Vegetation Science, 20:596–602
  • Zelený D. (2009): Co-occurrence based assessment of species habitat specialization is affected by the size of species pool: reply to Fridley et al. (2007). Journal of Ecology, 97:10–17
  • Zelený D. (2009): Jak se „botanizuje“ na Tchaj-wanu. Ekolist 4


  • Zelený D. & Li C.-F. (2008): Vegetace suchých acidofilních trávníků v oblasti třebíčského žulosyenitového plutonu [Dry acidophilous grassland vegetation in the area of Třebíč's granosyenite pluton]. Acta Rerum Naturalium, 5: 177–194
  • Zelený D. (2008): Lesní vegetace v údolí Vltavy severně od Zlaté Koruny (okres Český Krumlov) [Forest vegetation in the Vltava river valley north of Zlatá Koruna (district of Český Krumlov)]. Zprávy České Botanické Společnosti, 43: 111–169
  • Zelený D. (2008): 'Taiwan? A mají tam vůbec nějaký les?' ['Taiwan? And is there some forest?'] Živa 56 (1): 17–19


  • Zelený D. & Chytrý M. (2007): Environmental control of vegetation pattern in deep river valleys of the Bohemian Massif. Preslia, 79: 205–222
  • Hájková P., Hájek M., Apoštolová I., Zelený D., Dítě D. (2007): Shifts in the ecological behaviour of plant species between two distant regions: evidence from the base richness gradient in mires. Journal of Biogeography, 35: 282–294
  • Hájek M., Tichý L., Schamp B.S., Zelený D., Roleček J., Hájková P., Apostolová I. & Dítě D. (2007): Testing the Species Pool Hypothesis for mire vegetation: exploring the influence of pH specialists and habitat history. Oikos 116: 1311–1322
  • Boublík K. & Zelený D. (2007): Plant communities of silver fir forests in southeastern Bohemia. Tuexenia, 27: 73–90
  • Zelený D. (2007): Deep river valleys - one of the hot spots of biodiversity in Central Europe. Yearly Journal of Taiwan Biodiversity Association 2007 (into Chinese translated by Ching-Feng Li)


  • Zelený D. (2006): Výskyt vegetace širokolistých suchých trávníků svazu Bromion u Nové Vsi na Třebíčsku. [Occurrence of broad-leaved semi-dry grasslands of the Bromion alliance close to village Nová Ves, district of Třebíč (Czech Republic)] Acta Rerum Naturalium, Jihlava, 2: 39–44


  • Zelený D., Šraitová D., Mašková Z. & Květ J. (2001): Management effects on a mountain meadow plant community. Silva Gabreta, 7: 45–54